Darwinian Populations and Natural Selection by Godfrey-Smith Peter;
Author:Godfrey-Smith, Peter;
Language: eng
Format: epub
Publisher: Oxford University Press, Incorporated
Published: 2009-06-14T16:00:00+00:00
5.5. Summary of the First Five Chapters
Darwinian populations are collections of things that vary, reproduce at different rates, and inherit some of this variation. The basic features of these collections are startlingly routine—births, lives, and deaths, with variation and inheritance. But Darwin saw that this set-up, this arrangement of ordinary features, is an extraordinarily important element of the world. Darwin’s description was empirical and concrete. The last century’s work has included a series of moves towards abstraction, attempting to say what is essential about the Darwinian machine—which features are not dependent on the contingent particularities of life on earth. I continue that tradition, but do so with an eye to another feature of the Darwinian world view. Darwinian populations shade into marginal cases, and the paradigm Darwinian processes depend on ingredients that are themselves evolutionary products and must have come from something simpler. One aim of this book is to give an account of the Darwinian process that is designed to handle this blending-off into marginal cases, precursors, and not-quites.
This account of Darwinism yields a particular picture of the world. One of the world’s constituents is a great range of Darwinian populations: paradigm cases and marginal ones, some clear and others obscure, some powerful and others suppressed. Some are visible and obvious, others invisible. Some are inside others. They tread through their Darwinian behaviors on a great range of different scales in space and time. Some evolve via reproduction of a wholesale and definite kind, others evolve by coopting the biological scaffolding that results. Populations evolve as a consequence of their Darwinian properties, but also change the basis for their further evolution, moving through the imagined spaces of evolutionary parameters. The tree of life is generated by Darwinian populations and what they do—the tree is a structure of lives linked by reproductive events. But reproduction is an evolutionary product, and appears as a different relationship at different places on the tree. Sometimes there is sex, a fresh start, and genetic novelty with every birth; sometimes the appearance of a new organism is imperfectly distinguished from continuation of the same thing. Some Darwinian individuals live inside others, in ways that make it unclear how to count and distinguish them. And sometimes the tree shape is lost due to fusions and hybridizations.
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